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Table 2

List of all recorded NIMS in four major large Croatian rivers, with their origin and densities (min, average, max) per m2 within each river (*first record in Croatia; bold species: nine most widespread and abundant NIMS; adensity calculated without site DR3).

Taxa group Species Origin Recorded at rivers and sites: Density (ind. m−2)

Average (min–max)
Turbellaria Dendrocoelum romanodanubiale* Ponto-Caspian Drava (2): DR13, DR14 10 (5–14)
Oligochaeta Branchiura sowerbyi Southern Asia Drava (6): DR4, DR7, DR13, DR14, DR15, DR18 70 (10–207)
Sava (5): SA13, SA14, SA17, SA2O, SA21 5 (1–14)
Potamothrix moldaviensis * Ponto-Caspian Drava (1): DR13 5 (5–5)
Danube (1): DA3
Polychaeta Hypania invalida Ponto-Caspian Drava (3): DR15, DR16, DR17 24 (10–43)
Danube (3): DA1, DA3, DA4 3 (2–4)
Hirudinea Caspiobdella fadejewi * Ponto-Caspian Sava (2): SA6, SA9 9 (5–12)
Gastropoda Potamopyrgus antipodarum New Zealand Mura (1): MU2 10 (10–10)
Drava (11): DR3, DR4, D4, DR6, DR8, DR9, DR10, DR11, DR12, DR13, DR14 87a (4–127.762)
Physella acuta North America Drava (2): DR13, DR14 8 (5–15)
Sava (1): SA17 2 (2–2)
Danube (1): DA4 1 (1–1)
Bivalvia Corbicula fluminea East Asia Drava (8): DR13, DR14, DR15, DR16, DR17, DR18, DR19, DR20 282 (6–1084)
Sava (16): SA5, SA7, SA8, SA9, SA10, SA11, SA12, SA13, SA14, SA15, SA16, SA17, SA18, SA19, SA20, SA21 54 (1–559)
Danube (3): DA1, DA2, DA4 41 (6–91)
Dreissena polymorpha Ponto-Caspian Drava (17): DR1, DR4, DR6, DR7, DR8, DR9, DR10, DR11, DR12, DR13, DR14, DR15, DR16, DR17, DR18, DR19, DR20 424 (24–2496)
Danube (4): DA1, DA2, DA3, DA4 26 (6–75)
Dreissena bugensis Ponto-Caspian Danube (4): DA1, DA2, DA3, DA4 15 (2–52)
Sinanodonta woodiana East Asia Drava (1): DR18 3 (3–3)
Sava (2): SA6, SA10 2 (1–2)
Danube (1): DA2 2 (2–2)
Amphipoda Chelicorophium sowinskyi Ponto-Caspian Drava (1): DR18 11 (11–11)
Sava (14): SA8, SA9, SA10, SA11, SA12, SA13, SA14, SA15, SA16, SA17, SA18, SA19, SA20, SA21, 338 (4–1568)
Chelicorophium curvispinum Ponto-Caspian Drava (11): DR10, DR11, DR12, DR13, DR14, DR15, DR16, DR17, DR18, DR19, DR20 2607 (3–7964)
Sava (4): SA8, SA9, SA10, SA17 22 (7–50)
Danube (4): DA1, DA2, DA3, DA4 132 (95–191)
Chelicorophium robustum Ponto-Caspian Danube (3): DA1, DA2, DA3 18 (4–33)
Dikerogammarus bispinosus Ponto-Caspian Danube (1): DA2 4 (4–4)
Dikerogammarus haemobaphes Ponto-Caspian Sava (14): SA7, SA8, SA10, SA11, SA12, SA13, SA14, SA15, SA16, SA17, SA18, SA19, SA20, SA21 218 (3–626)
Danube (1): DA1 2 (2–2)
Dikerogammarus villosus Ponto-Caspian Drava (15): DR5, DR6, DR8, DR9, DR10, DR11, DR12, DR13, DR14, DR15, DR16, DR17, DR18, DR19, DR20 809 (16–3307)
Sava (3): SA8, SA9, SA15 47 (23–91)
Danube (4): DA1, DA2, DA3, DA4 128 (57–204)
Echinogammarus ischnus Ponto-Caspian Drava (6): DR14, DR15, DR16, DR17, DR19, DR20 93 (3–278)
Danube (3): DA1, DA3, DA4 6 (5–7)
Obesogammarus obesus Ponto-Caspian Danube (3): DA1, DA2, DA4 18 (6–36)
Isopoda Jaera istri Ponto-Caspian Drava (9): DR10, DR11, DR12, DR14, DR15, DR16, DR17, DR18, DR19 394 (7–1459)
Sava (12): SA8, SA9, SA10, SA11, SA12, SA14, SA16, SA17, SA18, SA19, SA20, SA21 72 (3–391)
Danube (3): DA1, DA3, DA4 81 (2–238)
Mysidacea Limnomysis benedeni Ponto-Caspian Drava (2): DR19, DR20 9 (1–16)
Danube (2): DA1, DA3 2 (1–2)

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