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Invertebrate bioassessment metrics over time from the Niobrara River at Agate Fossil Beds National Monument, Nebraska, USA. (a) Hilsenhoff's biotic index (HBI), (b) Ephemeroptera, Plecoptera and Trichoptera (EPT) richness, (c) taxon richness, and (d) proportion of EPT taxa. Vertical dashed lines indicate where past data (1989–2009) from Bowles et al. (2013) end and newly collected data for both HD (solid) and Hess (dashed) lines begin.
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