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Cited article:

From Late Miocene to Holocene: Processes of Differentiation within the Telestes Genus (Actinopterygii: Cyprinidae)

Vincent Dubut, Antoine Fouquet, Adrien Voisin, et al.
PLoS ONE 7 (3) e34423 (2012)
https://doi.org/10.1371/journal.pone.0034423