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Cited article:

Which is the major trigger in aquatic environment for pearl oyster Pinctada fucata martensii sperm from gonad: Ammonia ion or pH ?

Xing Zheng, Zhenyi Fu, Siqi Lin, et al.
Aquaculture 520 734673 (2020)
https://doi.org/10.1016/j.aquaculture.2019.734673

Motility activation and metabolism characteristics of spermatozoa of the black-lip-pearl oyster Pinctada margaritifera var: cumingii (Jameson, 1901)

M. Demoy-Schneider, A. Levêque, N. Schmitt, M. Le Pennec and J. Cosson
Theriogenology 77 (1) 53 (2012)
https://doi.org/10.1016/j.theriogenology.2011.07.014

The ageing phenomenon of turbot spermatozoa: effects on morphology, motility and concentration, intracellular ATP content, fertilization, and storage capacities

M. Suquet, C. Dreanno, G. Dorange, Y. Normant, L. Quemener, J. L. Gaignon and R. Billard
Journal of Fish Biology 52 (1) 31 (1998)
https://doi.org/10.1111/j.1095-8649.1998.tb01550.x