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Cited article:

General patterns of sexual dimorphism in graylings (Thymallus), with a comparison to other salmonid species

Gernot K. Englmaier, Alexander Antonov and Steven J. Weiss
Reviews in Fish Biology and Fisheries 32 (2) 645 (2022)
https://doi.org/10.1007/s11160-021-09694-4